词根词缀：acto-, act-, akto-, akt-

【来源及含义】Greek > Latin: seashore, beach, strand, promontory

【同源单词】Actiornis, Actitis macularia, actophile, actophilous, aktios, aktites

词根词缀：macul-, maculat-

【来源及含义】Latin: spot, mark, stain, blot, blemish, mesh; the original meaning of macula seems to have been, "a soiled spot, a spot to be cleaned"

【同源单词】Actitis macularia, immaculate, lepra maculosa, macula, macula cerulea, macula retina